Two civilian virologists, Drs. Rachel Scott and Quincy Tophet, claiming to be studying birds, join Captain Tom Chandler and crew aboard the USS Nathan James on their way to the Arctic.
Renegade Russian forces attack them, and Chandler learns the virologists are actually collecting samples of a deadly virus that has infected 80 percent of the human population while they have been at sea. The remaining U.S. government orders the ship home, and those aboard try desperately – many in vain – to contact their family. After a nuclear detonation in France derails their plans for refueling, the crew boards a ship filled with dead, infected people.
While originally setting sail for Naval Station Mayport, after losing contact with the government, Chandler decides that the best place to work on the vaccine is on the ship itself out in open sea. Dr. Tophet is revealed to be secretly communicating with the Russians.
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